male Canada goose is determined by the rank of his family. 9, the first known use in an educational context is in the. The national qualifications include the Life Skills Coursework (SFL National 3 (NAT3 National 4 (NAT4 National 5 (NAT5 Higher, and Advanced Higher. "Dominance, cortisol and stress in wild chimpanzees". London: Chapman and Hall, 1987 page needed Samuels,.; Silk,. This polygynous behavior has also been observed in some eusocial bees such as Schwarziana quadripunctata. "Egalitarian Behavior and Reverse Dominance Hierarchy". 7 In great tits and pied flycatchers, high-ranking individuals experience higher resting metabolic rates and therefore need to consume more food in order to maintain fitness and activity levels compared to subordinates in their groups. Former research suggests that primer pheromones secreted by the queen cause direct suppression of these vital reproductive hormones and functions however current evidence suggests that it is not the secretion of pheromones which act to suppress reproductive function but rather the queen's extremely high levels. In populations of Kenyan vervets, high-ranking females have higher foraging success when the food resources are clumped, but when food is distributed throughout an area they lose their advantage because subordinate females can acquire food with less risk of encountering a dominant female. The ecology of relationships.